Elderberries are quite edible. The blue or purple
ELDERBERRY
berries are gathered and made into elderberry wine, jam, syrup, and pies. The entire flower cluster can be
Sambucus nigra L. ssp.
dipped in batter and fried while petals can be eaten raw or made into a fragrant and tasty tea. The flowers
canadensis (L.) R. Bolli
add an aromatic flavor and lightness to pancakes or
The elderberry is of well-known value to the Indians
Contributed by: USDA NRCS National Plant Data
of North America and the many purposes it serves
Center & the Biota of North America Program
(Barrow 1967). Elderberry is highly prized by both Spaniards and Cahuillas. Throughout the months of July and August the small clusters of berries are gathered in large quantities. These clusters are dried carefully on the drying floor and preserved in considerable amounts. When wanted, they are cooked into a rich sauce that needs no sweetening. A Cahuilla family during this season of the year will subsist largely on these messes of "sauco." Frequently, the elderberry was so greatly enjoyed that families would live for weeks on little else. Many were dried for use in the winter, and were either re-
Botany Dept., NMNH, Smithsonian Institution
cooked or eaten raw. Elderberries are still highly
prized for food by modern Indian people.
Alternate Names
Elderberry twigs and fruit are employed in creating
Arizona elderberry, American elder, sweet elder, wild
dyes for basketry. These stems are dyed a very deep
elder, flor sauco, tree of music, Danewort, Walewort,
black by soaking them for a week or so in a wash
New Mexican elderberry, velvet-leaf elder, hairy blue
made from the berry stems of the elderberry
elderberry, and dwarf elder. Taxonomically, there
(Barrows 1967). The Cahuilla split basketry
have been recent changes in this elderberry species.
materials from the aromatic sumac (Rhus trilobata).
It was previously divided into Sambucus coriacea,
Sambucus orbiculata, Sambucus velutina, and
Elderberry branches were used to make the shaft of
Sambucus caerulea (Munz 1968). This species is
arrows. Flutes and whistles were constructed by
known in some floras as Sambucus mexicana.
boring holes into stems hollowed out with hot sticks.
Clapper sticks were made by splitting the stem and
clapping the two halves against each other. Clapper
Ethnobotanic: Only the blue or purple berries of
sticks were used ceremonially in the round-house to
elderberry are edible. Edible berries and flower are
accompany singing and dancing. The pith of the
used for medicine, dyes for basketry, arrow shafts,
stems was used as tinder, and the stem itself was
flute, whistles, clapper sticks, and folk medicine.
employed as a twirling stick for starting the fire.
The active alkaloids in elderberry plants are
Hollowed-out elderberry stems can be made into
hydrocyanic acid and sambucine. Both alkaloids will
cause nausea so care should be observed with this
plant. Elderberries are high in Vitamin C. The red
In the middle ages elderberry was considered a Holy
berries of other species are toxic and should not be
Tree, capable of restoring good health, keeping good
The wood is hard and has been used for combs,
Fruits of elderberry are gathered from the wild for
spindles, and pegs, and the hollow stems have been
wine, jellies, candy, pies, and sauces. The plants are
commercially cultivated for fruit production in
Plant Materials <http://plant-materials.nrcs.usda.gov/> Plant Fact Sheet/Guide Coordination Page <http://plant-materials.nrcs.usda.gov/intranet/pfs.html> National Plant Data Center <http://npdc.usda.gov>
Oregon. Sambucus canadensis and S. nigra have
create overhanging banks, providing habitat for fish
long been used in the same way, and cultivars of both
have been developed. All parts of the elderberry
plant are considered to be a valuable healing plant in
Game birds, squirrels and other rodents, and several
many folk medicine traditions (Hutchens 1991,
kinds of browsers also feed on the fruit or foliage of
Walker et al. 1993; Barrett et al. 1933; Clarke 1977).
elderberry. Bears love to eat the elderberry fruits
Elderberry flowers contain flavenoids and rutin,
while deer, elk, and moose browse on the stems and
which are known to improve immune function,
foliage. The elderberries are important sources of
particularly in combination with vitamin “C.” The
summer food for many kinds of songbirds. For
flowers also contain tannins, which account for its
example, the western bluebird, indigo bunting,
traditional use to reduce bleeding, diarrhea, and
common house finch, red-shafted flicker, ash-
throated flycatcher, black-headed grosbeak, scrub
jay, Stellar jay, ruby-crowned kinglet, mockingbird,
The flowers are the mildest part of the plant and
red-breasted nuthatch, Bullock’s oriole, hooded
prepared as a tea, are used to break dry fevers and
oriole, song sparrow, white-crowned sparrow,
stimulate perspiration, aid headache, indigestion,
western tanager, California thrasher, russet-backed
twitching eyes, dropsy, rheumatism, appendix
thrush, brown towhee, Audubon warbler, cedar
inflammation, bladder or kidney infections, colds,
waxwing, Lewis and Nuttall's woodpecker, wren-tit,
influenza, consumption (bleeding in lungs), and is
grouse, pheasant, and pigeons all eat elderberries
helpful to newborn babies (Hutchens 1991). Used as
a wash, the flowers or leaves are good for wounds,
sprains, and bruises, as well as for sores on domestic
The valley elderberry longhorn beetle (VELB)
animals. The leaves, which are stronger, have a
(Desmocerus californicus dimorphus) was listed as
slightly laxative property. Applied externally, leaves,
threatened under The Endangered Species Act on
flowers, bark and twigs are excellent as a poultice,
August 8, 1980. The elderberry beetle is endemic to
mixed equally with chamomile, for soreness,
moist valley oak riparian woodlands along the
inflammations, joint stiffness, and to reduce the
margins of rivers and streams in the lower
swelling of bee stings. The flowers and berries,
Sacramento and upper San Joaquin Valley of
employed as a diuretic, can aid arthritis and
California where elderberry grows. The primary
rheumatism. Steeped in water, the flowers are used
threat to the VELB is loss of habitat, insecticide and
externally to aid in complexion beauty, tone and
herbicide use, and lack of elderberry shrubs/trees as a
soften the skin, and lighten freckles or spots. The
food plant for the beetle. The mitigation for VELB
berry juice made into salve aids burns and scalds.
habitat loss, considered a taking under The
The juice taken internally will act as a purgative.
Endangered Species Act, is quite stringent (U.S. Fish
and Wildlife Service Mitigation Guidelines).
Livestock: Elderberry is a useful range plant for
domestic livestock, but is not equally palatable
In general, longhorn beetles are characterized by
during all seasons. It is usually receives limited
somewhat elongate and cylindrical bodies with long
browsing in the spring and to a much greater extent
antennae, often in excess of 2/3 of the body length.
in the late summer and fall. The leaves are eagerly
Male VELB have a metallic-green pattern of 4
devoured after the first heavy frost in the fall.
oblong maculations, surrounded by a bright red-
Because many branches are beyond the reach of the
orange border. The body length is about 13-21 mm,
animals, utilization is less destructive. Browse
and antennae are about a long as the body. Females
rating: Good for goats; good to fair for sheep; good to
are more robust, with body length about 18-25 mm,
poor for deer; fair for cattle; and fair to poor for
Elderberry is planted because of its forage and cover
Wildlife: Structurally complex riparian vegetation
value, productivity, adaptability, and ease of
communities provide many different habitats and
establishment. It is a useful ground cover for
support a diverse array of animal species. Different
stabilizing streambanks and eroding sites. It provides
groups of animals occupy or use the different layers
food, cover, perching, and nesting sites for many
of vegetation, and this multi-story arrangement is
species of birds and food and cover for various other
often present nowhere else in the arid landscapes.
wildlife, and it is important as browse for mule deer
Canopies of plants growing on stream banks provide
and elk. In the spring the leaves may be strongly
shade, cooling stream water, while roots stabilize and
scented and less palatable, but they sweeten and become more palatable by fall.
Adaptation
Elderberry grows on moist, well-drained sunny sites,
Please consult the PLANTS Web site and your State
usually occurring in early seral communities or in
Department of Natural Resources for this plant’s
openings in moist forest habitats (slopes, canyons,
current status, such as, wetland indicator values.
cliff bases, streamsides, streambanks) and moist areas
Western riparian ecosystems have been greatly
within drier, more open habitats (sagebrush,
altered by human activity. Riparian forests have been
mountain brush, pinyon-juniper, ponderosa pine,
reduced to fragmented, discontinuous patches
often along fence rows and roads); at elevations of 3-
because of human intervention. For example,
3000 meters. Elderberry is a dominant understory
estimates are that 70 - 90 percent of the natural
species in riparian woodlands. It can persist past
riparian ecosystems in the U.S. have been lost to
seral stages as scattered individuals in open forests,
human activities. Regional losses in these
woodlands, chaparral, or riparian zones. This species
ecosystems have been estimated to exceed 98% in the
flowers from May to September and fruits from July
Sacramento Valley in California. Many factors have
to October. Common elderberry is more common on
contributed to these resource losses, including the
warmer sites than red elderberry (Sambucus
following: natural resource use; urbanization;
racemosa), although they overlap in habitat
alteration of stream flows through dam construction
and ground-water withdrawal; modification of biotic
conditions through grazing, agriculture, introduction
In California, common riparian woodland associates
of non-native species; and alteration within
are valley oak (Quercus lobata), interior live oak
(Quercus wislizenii), California walnut (Juglans hindsii), and California sycamore (Platanus Description racemosa). Box elder (Acer negundo), Oregon ash
General: Honeysuckle family (Caprifoliaceae).
(Fraxinus latifolia), alder (Alnus rhombifolia), and
Native shrubs growing 2-4(-8) m tall, less commonly
willow (Salix gooddingii, Salix exigua, Salix
small single-stemmed trees, young twigs soft and
lasiandra, and Salix laevigata) are particularly
pithy but the wood hard; bark thin, grayish to dark
prevalent in the subcanopy. Understory species are
brown, irregularly furrowed and ridged. The
mostly shrubs, including elderberry (Sambucus
pinnately compound leaves are deciduous, opposite,
mexicana), buttonbush (Cephalanthus occidentalis),
about 15-35 cm long, odd-pinnate with (3-)5-9
blackberry (Rubus spp.), and California rose (Rosa
serrate leaflets 2-15 cm long, often with a long stalk,
californica). Lianas, such as poison oak
often asymmetrical at the base. Elderberry leaves,
(Toxicodendron diversiloba) and California grape
especially on seedlings or shrub-sized plants (without
(Vitis californica) are a dominant feature.
fruits or flowers) resemble California walnut
Herbaceous vegetation is 1% cover except in
(Juglans hindsii) and Oregon ash (Fraxinus latifolia).
The inflorescence is flat-topped, 4-20(-30) cm across,
broader than high; flowers bisexual, the corollas
Establishment
small, white to cream, rotate, 5-lobed, with a
Elderberry produces a good seed crop almost every
pleasant, yet slightly rancid odor. Fruit is berry-like,
year. The seeds are dispersed by birds and other
5-6 mm wide, with 3-5 nutlets, blue- to purple-black
animals that eat the fruit. The seeds have a hard seed
at maturity with a white-waxy bloom and appearing
coat and embryo dormancy and may remain viable
powder blue. The common name “elder” is from the
for up to 16 years in storage. Without pretreatment,
Anglo-Saxon “ellen,” meaning fire-kindler, the dry,
seed germination may be delayed from 2 to 5 years
after planting. Plants may flower and fruit after only
2-3 years and can reach full size in 3-4 years. They
Distribution: Common elderberry is common along
are said to be “short-lived.” Vegetative reproduction
stream banks, river banks, and open places in riparian
is limited to coppicing if the stems are killed or
areas lower than < 3000 m. From west Texas north
to Montana, western Alberta, and southern British
Columbia, and all other western states, south into
northwest Mexico. For current distribution, please
consult the Plant Profile page for this species on the
Elderberry fruits are collected when ripe and spread
To separate seeds from fruits either 1) run fruit
through a macerator with water and the pulp and empty seeds float; 2) crush, dry and use without
separating fruits and seeds; and 3) for small
• Cut tree back to 3 to 6 feet from the ground or to
amounts of fruit they can be cleaned in a fruit
50 percent of its height (whichever is greater) by
removing branches and stems above this height.
Elderberry seed can be stored dry at 41 °F for
The trunk and all stems greater than 1.5 inches in
Elderberry seeds can be sown in the fall soon
• If the presence of tunnels excavated by the beetle
after collection, or stratified and sown in the
larvae inside the elderberry stem and trunk are
spring. In either case, germination is often not
A seedling density of 35 plant per square foot is
• Move plant by the root ball. Keep the root ball
sought. Seeds are sown 1/4 inch deep in drills
and covered with about 3/8 inch of sawdust
• Place transplant in holes 3 to 4 feet deep.
• Construct circular water retention basin from the
In the greenhouse, seeds are warm stratified for
excavated earth about 8-10 feet in diameter and
two months in a mixture of peat, vermiculite, and
12 to 14 inches high. Plant the main trunk of an
sand at 21° to 30°C; (70 to 85°F). Seeds are then
elderberry in the center of each water basin.
placed close to the soil surface in flats in the
Plant other stems that have been rooted around
greenhouse. There are usually several hundred
• Saturate soil with water. Irrigate as necessary,
Seedlings are then potted from the flats in deep
especially through first growing season.
After one season of growth, the seedlings are
Management
field planted in the fall or spring when they are 6
In six riparian restoration projects carried out in
to 8 months old. If planted in the fall irrigation
California, competition from exotic weed species was
may not be necessary in moist sites. In drier
a key factor in mortality and site failure (Baird 1989).
sites or with spring planting, irrigation will be
On small sites, hand weeding around trees and shrubs
is the most effective means of weed control. One
way to avoid competition from weeds on larger sites
is to remove the surface soil, although this has the
Cuttings of elderberry tend to have lower
disadvantage of removing nutrients, mycorrhizal
survival success than establishment from seed.
fungi, bacteria, and insect and invertebrate
Use hardwood cuttings from previous seasons
populations critical to a healthy habitat. A cover crop
of native wildflowers was also used to control weeds,
Take "heel cuttings" from older wood, so inner
with wildflower seeds hand-broadcast over the site.
On wetter, heavier soils this does not seem to provide
Cuttings should be at least 10", and have at least
two nodes. Cuttings are placed in 4" pots with
perlite and peat. Plants are kept moist.
There is considerable evidence that fertilization can
Cuttings have a fragile root system, with high
favor exotic weeds over native plants. Inoculation
mortality occurring when transplanted. Care
with mycorrhizal fungi enables seedlings of some
should be taken to be very gentle with delicate
species to better utilize limited supplies of both water
and nutrients. Inoculation of transplanted shrubs may
The cuttings, which do survive, seem to establish
be accomplished through inclusion of large (1.2 m
and grow faster. Plant biomass production,
deep by 2.8 m wide) root balls with plants. Smaller,
height, flowering and seed set is more rapid than
more economical soil plugs scattered throughout the
site serve the same purpose. The number of soil
plugs needed to ensure the establishment of soil flora
is directly related to the distance of the restoration
• All elderberry plants with evidence of valley
elderberry longhorn beetle use (i.e. emergence
Given that elderberry provides habitat for the
transplanted, as they provide habitat for a
federally listed valley longhorn elderberry beetle,
livestock grazing of elderberry is not recommended.
Species Act. For further technical information,
Livestock grazing can alter vegetative structure and
call a representative of the U.S. Fish and
composition of riparian habitat. Overgrazing by
livestock and big game frequently changes plant
species composition and growth form, density of
Bolli, R. 1994. Revision of the genusSambucus.
stands, vigor, seed production of plants, and insect
Burke, H.E. 1921. Biological notes on Desmocerus ,
Clear-cutting or seed tree cutting with high soil
a genus of roundhead borers, the species of which
disturbance sometimes favors the development of
infests various elders. J. Econ. Ent. 14:450-452.
elderberry in a seral community. It recovers well
from heavy grazing in the Great Basin. For use in
Clarke, C.B. 1977. Edible and useful plants of
site stabilization or rehabilitation, seeds may be
California. University of California Press. 280 pp.
planted directly or seedlings and 1-2-year old stock
may be transplanted. It also grows from transplanted
Crane, M. F. 1989. Sambucus cerulea. IN: W.C.
Fischer (compiler). The fire effects information system [Data base]. USDA, Forest Service,
Elderberry usually is not present in the understory of
Intermountain Research Station, Intermountain Fire
closed-canopy forests, and when fire occurs in these,
regeneration occurs from seed banks that may occur
<http://www.fs.fed.us/database/feis/>
between 2-10 cm deep in the soil, the seeds deposited
from off-site dispersal or from plants of an earlier
Cronquist, A., A.H. Holmgren, N.H. Holmgren et al.
community. Fire scarifies the hard seed coat of
1984. Intermountain flora: Vascular plants of the
buried seeds and stimulates their germination, which
Intermountain West, U.S.A. Vol. 4. Subclass
usually occurs the first growing season after the fire.
Asteridae, (except Asteraceae). The New York
Subsequent burns may eliminate elderberry since it
spreads slowly by seed. Fire kills above-ground parts
but the root crown may sprout but a severe fire can
Eriksson, T. & M.J. Donoghue 1997. Phylogenetic
kill the root and stem buds from which sprouting
relationships of Sambucus and Adoxa (Adoxoideae,
Adoxaceae) based on nuclear ribosomal ITS
sequences and preliminary morphological data. Syst.
Cultivars, Improved and Selected Materials (and area of origin)
These plant materials are somewhat available from
Felter, H.W. & J.U. Lloyd 2000. Sambucus. IN
commercial sources. Contact your local Natural
King’s American Dispensatory. Web site.
Resources Conservation Service (formerly Soil
<http://metalab.unc.edu/herbmed/eclectic/kings/samb
Conservation Service) office for more information.
Look in the phone book under ”United States
Government.” The Natural Resources Conservation
Finn, C. 1999. Temperate berry crops. Pp. 324–334.
Service will be listed under the subheading
IN: J. Janick (ed.), Perspectives on new crops and new uses. ASHS Press, Alexandria, Virginia.
<http://www.hort.purdue.edu/newcrop/proceedings19
References
Arnold, R.A. 1984. Interim report for contract C-616 with the California Department of Fish and
Hartmann, H.T., D.E. Kesler, & F.T. Davies, Jr.
1990. Plant propagation principles and practices.
Prentice Hall, Englewood Cliffs, New Jersey. 647 pp.
Baird, K. 1989. High quality restoration of riparian ecosystems. Restoration and Management Notes
Hickman, J.C. (ed.) 1993. The Jepson manual. Higher plants of California. University of California
Barrett, S.A. & E.W. Gifford 1933. Miwok material culture. Indian Life of the Yosemite Region.
Hutchens, A.R. 1991. Indian herbalogy of North
Yosemite Association, Yosemite National Park,
America. Shambhala Books, Boston and London. pp
Barrows, D.P. 1967. Ethno-botany of the Coahuilla
Martin, A.C., H.S. Zim, and A.L. Nelson 1951.
Indians. Malki Museum Press. Banning, California.
American wildlife and plants: A guide to wildlife food habits. Dover Publications, Inc., New York, New
who require alternative means for communication of program information (Braille, large print, audiotape, etc.) should contact USDA's 720-2600 (voice and TDD).
Munz, P.A. 1968. A California flora. University of
To file a complaint of discrimination write USDA, Director, Office of Civil Rights, Room 326-W, Whitten Building, 14th and Independence Avenue, SW, Washington, DC 20250-9410 or call 202-720-5964 (voice or TDD). USDA is an equal opportunity
Murphy, E.V.A. 1959. Indian uses of native plants.
Mendocino County Historical Society. 81 pp.
Powers, S. 1976. Tribes of California. University of
Roos-Collins, M. 1990. The flavors of home. A guide to wild edible plants of the San Francisco Bay area. Heyday Books, Berkeley, California. 224 pp. Sampson, A.S. & B.S. Jesperson 1981. California range brushlands and browse plants. Agricultural Sciences Publications. University of California. Berkeley, California. 162 pp. Schopmeyer, C.S. 1974. Seeds of woody plants in the United States. Agriculture Handbook No. 450. Forest Service, U.S. Department of Agriculture. Washington, D.C. 883 pp. Walker, P.L. & T. Hudson 1993. Chumash healing. Changing Health and Medical Practices in an American Indian Society. Malki Museum Press, Banning, California. 161 pp. Prepared By Michelle Stevens Formerly USDA, NRCS, National Plant Data Center Guy Nesom Formerly BONAP, North Carolina Botanical Garden, University of North Carolina, Chapel Hill, North Carolina Species Coordinator M. Kat Anderson USDA, NRCS, National Plant Data Center, c/o Plant Sciences Department, University of California, Davis, California
Edited 03apr01 jsp; 03jun03 ahv; 060816 jsp For more information about this and other plants, please contact your local NRCS field office or Conservation District, and visit the PLANTS Web site Materials Program Web >
The U.S. Department of Agriculture (USDA) prohibits discrimination in all its programs and activities on the basis of race, color, national origin, sex, religion, age, disability, political beliefs, sexual orientation, and marital or family status. (Not all prohibited bases apply to all programs.) Persons with disabilities
WHAT IS LUPUS? patients 20:1. The disease is found most Lupus is an autoimmune disease which can affect the joints and muscles, It is characterized by varying degrees of inflammation in all or any of these sites. local fluid and pain. In Lupus, the injury antibodies, which, instead of protecting WHAT HAPPENS IN SLE? substances normally present in the body. weakness, fatigue, or
International Journal of Generic Drugs ANALYTICAL METHOD AM-V-0206-0300 P R O C E D U R E S of Pages: 24 HPLC ASSAY with DETERMINATION OF META-FLUOXETINE HCl. ANALYTICAL METHOD VALIDATION 10 and 20mg Fluoxetine Capsules HPLC DeterminationMethod is suitable for:ý In-process controlþ Product Releaseþ Stability indicating analysis (Suitability - US/EU Product)CAUTIONFLUOXETI